Proleg

In all the orders in which they appear, mainly Hymenoptera and Lepidoptera, prolegs of any form evolved independently of each other by convergent evolution. They appear to have evolved from ancient inactive genes that have since been activated again.

Prolegs of lepidopteran larvae have a small circle of gripping hooks, called "crochets". The arrangement of the crochets can be helpful in identification to family level. Although the point has been debated, prolegs are not widely regarded as true legs, derived from the primitive uniramous limbs. Certainly in their morphology they are not jointed, and so lack the five segments (coxa, trochanter, femur, tibia, tarsus) of thoracic insect legs. Prolegs do have limited musculature, but much of their movement is hydraulically powered.

Additional (stubby) prolegs on segment 8 have been reported on the Geometridae species Campaea perlata and Alsophila pometaria.